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Eventually, she learns to reconcile her actions after learning the truth about Innominat, and becomes more determined to defeat Artorius in order to make a better world free of 'reason'. Despite this however, Velvet still has her old feelings hidden inside of her, and is shown to secretly care for her comrades, especially a young malak whom she named Laphicet because he resembles her deceased brother. She has no qualms about killing anyone who stands in her way, and has racked up a large body count of daemons, malakhim, and exorcists alike. Velvet is shown to act very cruel, cynical, and only cares about her goal. Determined to avenge her brother's death, Velvet sets out on her long quest for revenge.

One day, a malak named Seres sets Velvet free from her prison to help her get revenge on Artorius, who has now formed an authoritarian society called the Abbey. Artorius quickly takes her down, and locks her up in the prison island of Titania for three years. She kills all of the surrounding daemons in a blind rage, but quickly finds out that the daemons she has killed are the villagers that Velvet knew. Laphicet was offered as a sacrifice to the fifth Empyrean Innominat, while Velvet was expelled out of the shrine with a new daemon arm, having been transformed into a Therion. Devastated, she tries to save her brother from falling into the shrine, but Artorius cuts off her left arm, causing both to fall into the shrine's depths. Worried about Laphicet's safety, Velvet rushes outside the village to a nearby shrine, where she finds Artorius killing her brother in some kind of sacrificial ritual. However, on the Scarlet Night, a massive outbreak of daemonblight turns all of the villagers into daemons. Velvet was generally optimistic, and deeply cared for the people around her, especially her brother Laphicet. Artorius, Celica's husband, was tasked with protecting the village and his family from daemons, and he taught Velvet everything he knew about his maxims to apply in combat.

She and her brother were the only Crowe siblings left after their older sister, Celica Crowe, was killed by daemons during a Scarlet Night three years prior. Velvet originally lived in the village of Aball, where she primarily hunted prickleboars for a living, and was taking care of her ill little brother Laphicet Crowe.

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